(((SIMEL1,(MESOR1,(BRMEL1,BRSUI1))),(RODEN1,(SILIC1,((SIPOM1,(RHSPH1,RHSPH2)),PADEN1)))),(CHSAL1,(((PHLUM1,(ERCAR1,(((SATYP1,(SATYI1,(SAENT4,(SAENT2,SAENT3)))),(SHSON1,(SHDYS1,(ESCOL6,(ESCOL9,(ESCOL5,(SHFLE3,(SHFLE2,(SHFLE1,(ESCOL2,ESCOL7)))))))))),(YEENT1,(YEPSE1,(YEPES5,(YEPES4,(YEPES3,(YEPES1,YEPES2))))))))),(ACPLE1,HADUC1)),((AEHYD1,(SHAMA1,((SHDEN1,SHFRI1),((SHEWA3,(SHEWA1,SHEWA2)),(SHONE1,SHEWA4))))),COPSY1))));