(((HADUC1,ACPLE1),(CHSAL1,(AEHYD1,((SHAMA1,(SHFRI1,(SHDEN1,((SHEWA3,(SHEWA2,(SHONE1,SHEWA1))),SHEWA4)))),COPSY1)))),(((SATYP1,(SATYI1,(SAENT4,SAENT3))),(SAENT2,((SHSON1,((SHDYS1,SHFLE3),(SHFLE2,(SHFLE1,(ESCOL9,(ESCOL7,(ESCOL6,(ESCOL5,ESCOL2)))))))),(((PHLUM1,(YEPSE1,YEENT1)),(YEPES4,(YEPES3,(YEPES2,(YEPES1,YEPES5))))),ERCAR1)))),((BRSUI1,(BRMEL1,(MESOR1,SIMEL1))),(RODEN1,((SIPOM1,SILIC1),(PADEN1,(RHSPH2,RHSPH1)))))));