((((PADEN1,(RHSPH1,RHSPH2)),(RODEN1,(SIPOM1,SILIC1))),((BRSUI1,BRMEL1),(SIMEL1,MESOR1))),(CHSAL1,((AEHYD1,((SHAMA1,((((SHEWA3,(SHEWA1,SHEWA2)),SHONE1),SHEWA4),(SHDEN1,SHFRI1))),(COPSY1,(ACPLE1,HADUC1)))),((((SATYI1,(SAENT3,SAENT4)),(SATYP1,SAENT2)),(ESCOL2,((SHFLE3,(SHFLE2,SHDYS1)),(SHSON1,((ESCOL7,(ESCOL9,(ESCOL6,ESCOL5))),SHFLE1))))),(ERCAR1,((YEENT1,(YEPSE1,(YEPES5,(YEPES4,(YEPES3,(YEPES1,YEPES2)))))),PHLUM1))))));